You're right that it can't be box 5, because that is the amount of after-tax dollars that you (not your employer) put into the 401(k). In fact, this makes me wonder why there is a number in box 5 at all.
I was going to say box 1 (the distribution), but it depends on what the person at Fidelity was thinking when they issued this form.
So contact Fidelity and ask them, "does box 1 contain only the original contribution or does it contain the contribution plus its earnings?" By the way, which amount is larger, boxes 1, 2a, or 5? Can you tell us the amounts?
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