Box (b) is the date you acquired the stock. It is significant to determine if the gain or loss on sale of the stock is long term (more than on year) or short term (less than on year). If you don't know exactly when the investment was acquired, you can pick a date the will result in the correct time period of one year or more for the amount of time you owned it.
Box (e) is what you paid to acquire the stock. If you don't know what that was, you can enter 0 for the entry, but that will probably result in a higher tax than if you enter the actual cost basis.
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